BIOS 102. Biological Sciences First-Year Seminar. 1 hour.

Seminar course designed to introduce first-year students in the Biological Sciences major to departmental resources and opportunities that will promote student success. Course Information: Satisfactory/Unsatisfactory grading only. Prerequisite(s): Open only to freshmen.

BIOS 104. Biology for Non-majors. 4 hours.

Origin and diversity of life; genetics, evolution, ecology and ecosystems; energy flow; photosynthesis; human anatomy and physiology; development of biological ideas; and biology, biotechnology and human society. Course Information: Credit is not given for BIOS 104 if the student has credit in BIOS 100 or BIOS 101 or BIOS 110 or BIOS 120. Animals may be used in instruction. Class Schedule Information: To be properly registered, students must enroll in one Laboratory-Discussion and one Lecture. Natural World - With Lab course.

BIOS 110. Biology of Cells and Organisms. 4 hours.

Introductory biology at the molecular, cellular, and organismal level. Topics include: Scientific skills, biological chemistry, cell structure and function, metabolism, cell division, molecular genetics, diversity, anatomy and physiology. Course Information: Previously listed as BIOS 100. Animals used in instruction. THIS COURSE IS INTENDED FOR SCIENCE MAJORS. BIOS 110 and BIOS 120 may be taken in any order. Credit is not given for BIOS 110 if the student has credit in BIOS 100. Credit is not given for BIOS 104 if the student has credit in BIOS 100, BIOS 101, BIOS 110 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Lecture and one Laboratory-Discussion. Natural World - With Lab course.

BIOS 112. Program Design I in the Context of Biological Problems. 3 hours.

Introduction to programming using Biology as the context for programming in a high-level language; control structures, variables, simple and aggregate data types; problem-solving techniques; biology topics include central dogma and genetics. Course Information: Same as CS 112. Credit is not given for BIOS 112 if the student has credit in CS 107 or CS 109 or CS 111 or CS 113. Course Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture-Discussion.

BIOS 120. Biology of Populations and Communities. 4 hours.

Introductory biology at the level of populations and communities. Topics include: Scientific skills, evolution, Mendelian and population genetics, biological diversity, and ecological systems including ecosystem processes and human impacts. Course Information: Previously listed as BIOS 101. Animals used in instruction. This course is intended for science majors. BIOS 110 and BIOS 120 may be taken in any order. Credit is not given for BIOS 120 if the student has credit in BIOS 101. Credit is not given for BIOS 104 if the student has credit in BIOS 100, BIOS 101, BIOS 110 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Lecture-Discussion and one Laboratory-Discussion. Natural World - With Lab course.

BIOS 184. The Basics of Neuroscience. 1 hour.

Introduction to the scientific study of the brain and behavior. Overview of neuroscience as an integrative discipline. Course Information: Same as PHIL 184 and PSCH 184. Satisfactory/Unsatisfactory grading only.

BIOS 196. Biology Colloquium. 2 hours.

Combines in-class seminars with off-campus day and weekend field trips, with the goal of introducing students to different areas of the Biological Sciences, and alternate career options.Course Information: May be repeated up to 1 time(s). Field trips required at a nominal fee. Prerequisite(s): Major in biological sciences, biochemistry, or neuroscience.

BIOS 199. Introduction to Research. 1 hour.

An introduction to research for students interested in becoming biology majors. Credit is contingent on approval by the research supervisor of a written report that is also submitted to the Biology colloquium faculty adviser. Course Information: 1 hour. Satisfactory/Unsatisfactory grading only. May be repeated. Prerequisite(s): Approval of the department. Class Schedule Information: To be properly registered, students must enroll in one Conference and one Laboratory.

BIOS 220. Genetics. 3 hours.

Principles of genetics, gene and chromosome structure, gene expression, inheritance of complex traits, Mendelian and non-Mendelian inheritance, genomes, and genetic engineering. Course Information: Prerequisite(s): BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Lecture-Discussion and one Lecture.

BIOS 222. Cell Biology. 3 hours.

The physiological processes that govern cell function. Topics include enzyme kinetics, metabolism, membranes, membrane transport, electrical and chemical signaling, intracellular motors, motility, and junctions. Prerequisite(s): BIOS 100 or BIOS 110.

BIOS 230. Evolution and Ecology. 3 hours.

Molecular and ecological basis of evolutionary change through adaptation or natural selection to explain diversity of species, behaviors, populations, communities and ecosystems and how these interactions are affected by the changing environment. Prerequisite(s): BIOS 101 or BIOS 120.

BIOS 236. Animal Behavior. 3 hours.

Examine the proximate and ultimate causes of animal behavior; neural and hormonal mechanisms; diversity of behavior and their relationship to an organism's ecology and evolution. Course Information: Prerequisite(s): BIOS 101 or BIOS 120.

BIOS 237. The Human Skeleton. 4 hours.

Examination of the human skeleton, emphasizing bone identification and functional anatomy. Course Information: Same as ANTH 237. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture.

BIOS 250. Microbiology and Application to Health. 3 hours.

Introduction to microbial growth and metabolism, disease pathogenicity and control, epidemiology and human infections. Course Information: Credit for BIOS 250 is not given if the student has credit in BIOS 350. Intended for pre-nursing students and should not be taken by those who need BIOS 350. Prerequisite(s): BIOS 100 or BIOS 110; and Credit or concurrent registration in CHEM 130 or CHEM 230 or CHEM 232 or CHEM 115. Recommended background: BIOS 101 or BIOS 120.

BIOS 272. Comparative Vertebrate Anatomy. 4 hours.

Comparative vertebrate anatomy through the analysis of varying morphological features of hard and soft tissues. Course Information: Animals used in instruction. Prerequisite(s): BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture.

BIOS 286. The Biology of the Brain. 3 hours.

Survey of basic neurobiology. Brain structure, chemistry, development and control of behavior (sensation, movement, emotions, memory, cognition, sex). Course Information: Prerequisite(s): BIOS 100 or BIOS 110.

BIOS 294. Special Topics in Biological Sciences. 1-5 hours.

Selected topics in the Biological Sciences. Course Information: May be repeated. Students may register for more than one section per term.

BIOS 296. Directed Instruction. 2 hours.

Leadership opportunity for upper-division students who wish to assist with the biology colloquium.Course Information: May be repeated to a maximum of 4 hours. Field trips required at a nominal fee. Students will be required to submit a final report describing their leadership activities and experiences. Prerequisite(s): BIOS 196 and approval of the department. To be properly registered, students must enroll in one Lecture-Discusssion and one Practice.

BIOS 299. Honors Biology. 1 hour.

An additional hour of related work for students registered in another course in Biological Sciences. Course Information: May be repeated. Prerequisite(s): Membership in Honors College, or, for superior students, approval of the department; and registration in a Biological Sciences course (except BIOS 391 or BIOS 399) and consent of the instructor. Open only to freshmen, sophomores, and juniors. Honors course.

BIOS 305. Plant Evolutionary Ecology. 3 hours.

Fundamental aspects of plant structure and function, plant-environment interactions, and how they shape biodiversity, with an emphasis on their evolutionary context. Prerequisite(s): BIOS 230.

BIOS 310. Genetics Laboratory. 2 hours.

Advanced laboratory techniques in gene inheritance, expression and regulation using genetic engineering, molecular biology and bioinformatic and statistical analysis to study Mendelian, microbial, molecular, human and population genetics. Course Information: Previously listed as BIOS 221. Animals used in instruction. Credit is not given for BIOS 310 if student has credit in BIOS 221. Prerequisite(s): Credit or concurrent registration in BIOS 220.

BIOS 312. Cell Biology Laboratory. 2 hours.

Laboratory training in advanced cell biology methods used in research and clinical settings. Microscopy, protein extraction and quantification, immunoassays, cell culture, experimental design, data analysis and scientific writing are emphasized. Course Information: Previously listed as BIOS 223. Credit is not given for BIOS 312 if the student has credit in BIOS 223. Prerequisite(s): Credit or concurrent credit in BIOS 222.

BIOS 320. Developmental Biology. 3 hours.

Principles governing growth and differentiation from the molecular to the organismic level. Course Information: Prerequisite(s): BIOS 220.

BIOS 321. Developmental Biology Laboratory. 3 hours.

Laboratory problems in developmental biology. Course Information: Animals used in instruction. Prerequisite(s): Credit or concurrent registration in BIOS 320.

BIOS 323. Molecular Biology Laboratory. 3 hours.

Modern molecular biology techniques used in the isolation and functional analysis of genes. Bioinformatics, gene cloning, gene fusions and expression assays will be used in the analysis of the gene isolation and analysis. Course Information: Prerequisite(s): BIOS 220.

BIOS 325. Human Embryology. 3 hours.

Major events in human development between fertilization and birth. The developmental origins of adult body structure and function, normal anatomical variations, and congenital malformations. Course Information: Prerequisite(s): BIOS 100 or BIOS 110.

BIOS 326. Embryology Laboratory. 1 hour.

Examination of histological sections of mammalian embryos. Identification of cells and tissues from their microscopic appearance, reading serial sections. Prerequisite(s): Credit or concurrent registration in BIOS 325.

BIOS 331. General Ecology Laboratory. 3 hours.

Field and laboratory data collection for hypothesis testing; required field trips to representative plant communities. Course Information: Animals used in instruction. Required field trips on Saturdays. Prerequisite(s): BIOS 101 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Practice.

BIOS 336. Animal Behavior Laboratory. 3 hours.

Field and laboratory data collection for hypothesis testing in animal behavior. Course Information: Animals used in instruction. Field trip required at a nominal fee. Field work required. Prerequisite(s): Credit or concurrent registration in BIOS 236.

BIOS 340. Environmental Physiology. 3 hours.

The performance of an organism or group of organisms in the context of their natural environment and their evolutionary background. The organism as an integrated system rather than a collection of cellular or organ processes. Course Information: Prerequisite(s): BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120.

BIOS 343. Animal Physiological Systems. 3 hours.

How the human body works including digestive, neuromuscular, sensory, respiratory, excretory, endocrine and cardiovascular systems using examples across animal species and human case studies. Course Information: Previously listed as BIOS 240. Credit is not given for BIOS 343 if student has credit in BIOS 240. Prerequisite(s): BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120. Recommended background: BIOS 222.

BIOS 350. General Microbiology. 3 hours.

An introduction to the morphology, staining, genetics, physiology, and biochemistry of microbial life with special emphasis on prokaryotic structure, function, diversity, and importance to human beings. Course Information: Prerequisite(s): BIOS 100 or BIOS 110; and Credit for CHEM 115 or Credit or concurrent registration in CHEM 130 or CHEM 230 or CHEM 232. Recommended background: BIOS 101 or BIOS 120.

BIOS 351. Microbiology Laboratory. 2 hours.

Laboratory experience with aseptic and pure culture techniques; staining and microscopy of microorganisms; identification of unknown bacteria; control of microbial growth; antibiotics; medical microbiology. Prerequisite(s): Credit or concurrent registration in BIOS 250 or BIOS 350.

BIOS 352. Introductory Biochemistry. 3 hours.

Structure and function of cellular constituents; enzymology; metabolism of carbohydrates, lipids, amino acids, nucleotides; molecular biology of biosynthesis of proteins and nucleic acids. Course Information: Same as CHEM 352. No credit given for BIOS 352 if student has credit in BIOS 452 or BIOS 454 or CHEM 452 or CHEM 454. No credit toward the degree in biochemistry. Prerequisite(s): BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120; and CHEM 230 or CHEM 232.

BIOS 360. Introduction to Paleontology. 4 hours.

The morphology, ecology, and relationships of fossil organisms. Basic principles of paleontology, including evolution, paleoecology and functional morphology. Course Information: Same as EAES 360. Field trip required at a nominal fee. One day field trip to collect fossils. Prerequisite(s): EAES 101 or EAES 111; or BIOS 100 or BIOS 110; and BIOS 101 or BIOS 120. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture.

BIOS 365. Human Ecological Systems. 3 hours.

Human-environmental interactions and feedbacks (positive and negative) and the ecological basis for health and sustainability of human-dominated landscapes. Case studies of real-world environmental issues. Course Information: Prerequisite(s): BIOS 101 or BIOS 120 or EAES 101 or PSCH 100 or SOC 100; or consent of the instructor.

BIOS 386. Seminar on Neurobiology. 2 hours.

Reading and discussion of both classic and recent research papers that are important in neurobiology. Course Information: Prerequisite(s): BIOS 286 or PSCH 262.

BIOS 391. Independent Study. 1 hour.

Individual study not covered in standard courses under close supervision of a faculty member. Credit is contingent on approval by research supervisor of a written report that is submitted to the department. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Students may register in more than one section per term. A maximum of 5 hours of BIOS 391 and/or BIOS 399 may be credited toward the department undergraduate major requirements. Prerequisite(s): Minimum of 2.00 grade point average in biological sciences courses and consent of the instructor. Class Schedule Information: This course counts toward the limited number of independent study hours accepted toward the degree and the major.

BIOS 399. Independent Research. 2 hours.

Individual research. Credit is contingent on approval by the research supervisor of a written report that is submitted to the department. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Students may register in more than one section per term. A maximum of 5 hours of BIOS 391 and/or BIOS 399 may be credited toward the department undergraduate major requirements. Prerequisite(s): Minimum of 2.00 grade point average in biological sciences courses, approval of the department, and consent of the instructor. Recommended background: Junior standing. Class Schedule Information: This course counts toward the limited number of independent study hours accepted toward the degree and the major.

BIOS 416. Natural Products. 3 or 4 hours.

Biogenetic approach to secondary metabolites. General principles and selected studies of phenolic compounds, terpenes, alkaloids, and other interesting natural products. Course Information: Same as CHEM 456. 3 undergraduate hours. 4 graduate hours. Prerequisite(s): One year of organic chemistry.

BIOS 420. Genomics. 3 hours.

Theoretical background in genomics with practical experience in obtaining and analyzing large scale Next Generation Sequencing (NGS) datasets using high capacity computational resources. Course Information: Extensive computer use required. Prerequisite(s): BIOS 220 or BIOS 230; or consent of the instructor. Recommended background: STAT 101 or STAT 130 or BIOS 112.

BIOS 427. Ecosystem Ecology. 3 or 4 hours.

Flow of energy and matter between the environment and biological organisms including bioological interactions that govern the cycling of water, carbon and nutrients at various spatial and temporal scales. Course Information: 3 undergraduate hours. 4 graduate hours. Prerequisite(s): BIOS 230.

BIOS 430. Evolution. 4 hours.

Focuses on empirical evolutionary data and theory, with an intro to data science, modeling, simulations in R programming language; it covers the history of evolution, population genetics, the species problem, biodiversity, macroevolution. Course Information: Extensive computer use required. Prerequisite(s): BIOS 120 AND either BIOS 220 or consent of the instructor. Recommended background: BIOS 230. Class Schedule Information: To be properly registered, students must enroll in one Lecture-Discussion and one Laboratory.

BIOS 431. Plant and Animal Interactions. 3 hours.

Ecology of plant and animal interactions. In-depth reading and discussion of primary literature on herbivory and plant defense and pollination, seed dispersal and protection mutualisms. Course Information: Prerequisite(s): BIOS 230.

BIOS 435. Plant Evolution. 3 hours.

Examines the history of plant life in a rigorous survey of plant genetics, factors that influence diversity of form and function, the astonishing diversity of plant sexual systems, and conservation. Course Information: Prerequisite(s): BIOS 230.

BIOS 437. Topics in Tropical Ecology. 3 hours.

Introduction to the character of tropical ecosystems. In-depth reading and discussion of one or more current topics. Course Information: Prerequisite(s): BIOS 230.

BIOS 443. Animal Physiological Systems Laboratory. 3 hours.

Discussion and laboratory exploration of mammalian physiological systems, including immune, endocrine, cardiac, vascular, nervous, pulmonary, renal, and digestive systems. Course Information: Animals used in instruction. Prerequisite(s): Credit or concurrent registration in BIOS 343; or Credit or concurrent registration in BIOS 340; or consent of the instructor. Class Schedule Information: To be properly registered, students must enroll in one Lecture, one Laboratory, and one Lecture-Discussion.

BIOS 446. Evolution and Human Disease. 3 hours.

Pathogen evolution, transmission of infectious disease, host response to pathogens, drug resistance in pathogens, and cancer progression/treatment are examined in light of natural selection, phylogenetics, coevolution, and population genetics. Course Information: Prerequisite(s): BIOS 230. Recommended background: BIOS 220.

BIOS 450. Advanced Microbiology. 3 hours.

Comprehensive analysis of metabolic, ecological, genomic, and functional diversity among the major groups of microorganisms. Relationship between microbial diversity and biogeochemistry in the environment, human/animal hosts, and engineered systems. Course Information: Prerequisite(s): BIOS 350.

BIOS 452. Biochemistry I. 4 hours.

Chemistry of proteins, nucleic acids, carbohydrates and lipids. Course Information: Same as CHEM 452. Prerequisite(s): Grade of C or better in CHEM 230; or Credit or concurrent registration in CHEM 234. Class Schedule Information: To be properly registered, students must enroll in one Discussion/Recitation and one Lecture.

BIOS 454. Biochemistry II. 4 hours.

Continues Biological Sciences 452. Carbohydrate and lipid metabolism, electron transport. Metabolism of amino acids, nucleic acids, proteins. Biosynthesis of macromolecules and regulation of macromolecular synthesis. Course Information: Same as CHEM 454. Prerequisite(s): BIOS 452 or CHEM 452. Class Schedule Information: To be properly registered, students must enroll in one Discussion/Recitation and one Lecture.

BIOS 455. Introduction to Landscape Ecology. 3 hours.

Uses a combination of lectures, discussions, and hands-on activities to introduce the concepts and methods of landscape ecology. We will discuss how landscape ecology can be applied to solving current environmental problems. Course Information: Extensive computer use required. Prerequisite(s): BIOS 230; or BIOS 331.

BIOS 458. Biotechnology and Drug Discovery. 3 or 4 hours.

Molecular and gene therapy, using small molecules including antisense, aptamers, and proteins. Structure-based drug design. Structural bioinformatics and drug discovery program. High-throughput screening. Combinatorial chemistry technology. Course Information: Same as CHEM 458. 3 undergraduate hours. 4 graduate hours. Prerequisite(s): BIOS 352 or CHEM 352; or Credit or concurrent registration in BIOS 452 or Credit or concurrent registration in CHEM 452; or consent of the instructor.

BIOS 466. Principles of Paleontology. 3 hours.

Theory and methods of evolutionary paleobiology; includes paleoecology, functional morphology, and major features of organic evolution. Course Information: Same as EAES 466. Prerequisite(s): EAES 360 or consent of the instructor. Class Schedule Information: To be properly registered, students must enroll in one Laboratory-Discussion and one Lecture.

BIOS 473. Soils and the Environment. 4 hours.

Soil science, emphasizing local soils and parent materials, soil classification and mapping, soil physics, soil gases and greenhouse gas emissions, soil chemistry and biogeochemistry, soil-plant interactions, and soil invertebrates. Course Information: Same as EAES 473. Field work required. Recommended background: Introductory courses in Chemistry and Biology are recommended. Coursework in EAES (such as EAES 101 and/or 111) is preferred.

BIOS 475. Neural Engineering I: Introduction to Hybrid Neural Systems. 3 or 4 hours.

Modeling and design of functional neural interfaces for in vivo and in vitro applications, electrodes and molecular coatings, neural prostheses and biopotential control of robotics. Course Information: Same as BME 475. 3 undergraduate hours. 4 graduate hours. Prerequisite(s): BIOE 472 or BME 472.

BIOS 480. Introduction to Modern Biostatistics with R. 3 hours.

An in-depth intro to research design, data visualization, and modern univariate statistics, from basic linear model to generalized linear models and linear mixed-effects models. Course Information: Extensive computer use required. All work done in the open-source R statistical computing language. Prerequisite(s): MATH 170 or MATH 180 or STAT 130. Class Schedule Information: To be properly registered, students must enroll in one Lecture-Discussion and one Laboratory.

BIOS 482. Molecular and Developmental Neurobiology Laboratory. 3 hours.

A hands-on laboratory course designed to explore the most recent neurotechniques and how they are being used to advance knowledge of the brain. Model organism use will be restricted to small invertebrates (C. elegans). Course Information: Prerequisite(s): BIOS 286; or PSCH 262.

BIOS 483. Neuroanatomy. 4 hours.

Organization of the nervous system, with an emphasis on mammals. Course Information: Same as PSCH 483 and NEUS 483. Animals used in instruction. Prerequisite(s): BIOS 272 or BIOS 286 or BIOS 325 or PSCH 262; or consent of the instructor. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture.

BIOS 484. Neuroscience I. 3 hours.

Neuroscience as an integrative discipline. Neuroanatomy of vertebrates, neural development, cellular neurobiology, action potential mechanisms, synaptic transmission and neuropharmacology. Course Information: Same as PHIL 484 and PSCH 484. Prerequisite(s): BIOS 286 or PSCH 262.

BIOS 485. Neuroscience II. 3 hours.

Integrative neuroscience, including sensory and motor systems; learning, memory, and language; pathology of nervous systems; philosophical perspectives, and modeling. Course Information: Same as PHIL 485 and PSCH 485. Prerequisite(s): BIOS 286 or PSCH 262.

BIOS 486. Animal Behavior and Neuroethology. 4 hours.

Neural and behavioral mechanisms of environmental information processing and interaction throughout the animal kingdom. Laboratory emphasizing a research project with journal style lab report, essay on assigned topic, occasional field trips required. Course Information: Animals used in instruction. Prerequisite(s): One course in neuroscience, animal physiology, or animal behavior. Class Schedule Information: To be properly registered, students must enroll in one Laboratory and one Lecture.

BIOS 489. Cellular Neurobiology Laboratory. 3 hours.

Recording from and analyzing the activity of nerve cells, neuronal networks, and other electrically excitable tissues. Course Information: Prerequisite(s): BIOS 286 or the equivalent.

BIOS 490. Topics Biological Sciences. 3 or 4 hours.

In-depth analysis of advanced topics in molecular, cellular, biomedical, ecology and evolution and neuroscience involving reading primary literature, term paper, student presentations and critical discussion. Course Information: 3 undergraduate hours. 4 graduate hours. May be repeated. Students may register in more than one section per term. Prerequisite(s): BIOS 230; or BIOS 220; or BIOS 222; or Graduate standing; or consent of the instructor.

BIOS 518. Geobiology. 4 hours.

Interactions between microorganisms and minerals, preservation of organisms and biofilms, influence of microorganisms in biogeochemical cycles, microorganisms on early Earth, life in extreme environments, the dark biosphere, and astrobiology. Course Information: Same as EAES 518. Recommended background: Basic knowledge of biology, chemistry, and earth sciences at the level of introductory college courses in each subject.

BIOS 520. Topics in Genetics. 2 hours.

Discussion of selected topics of current interest in genetics. Course Information: May be repeated. Students may register in more than one section per term. Prerequisite(s): BIOS 220 and BIOS 221 and consent of the instructor.

BIOS 523. Biology of MicroRNAs and other Small RNAs. 2 hours.

History, overview and biology of small RNA pathways, including microRNAs, siRNAs, RNA interference, roles in various biological processes, implication in disease pathophysiology, and potential therapies. Course Information: Same as ANAT 523. Prerequisite(s): Consent of the instructor.

BIOS 524. Molecular Biology Principles and Methods. 5 hours.

Guided reading and critical evaluation of foundational and current research topics in biochemistry and molecular biology. Course Information: Prerequisite(s): Prerequisite(s): Graduate standing; or consent of the instructor.

BIOS 525. Principles and Methods in Cell Biology. 5 hours.

Guided reading and critical evaluation of foundational and current research topics in cell biology. Course Information: Prerequisite(s): BIOS 524; and graduate standing; or consent of the instructor.

BIOS 526. Molecular and Genetic Analysis of Development. 3 hours.

Examines developmental mechanisms used in animal model systems. Course Information: Same as BCMG 526. Prerequisite(s): Graduate standing or consent of the instructor.

BIOS 527. Cellular and Systems Neurobiology. 3 hours.

Molecular and cellular properties of ion channels in neurons and sensory cells and their relationship to brain and sensory systems. Course Information: Same as ANAT 527 and NEUS 527. Prerequisite(s): Credit in one neuroscience course or consent of the instructor.

BIOS 528. Current Literature in Cell Biology. 3 hours.

Topics in cell biology. Course Information: Prerequisite(s): BIOS 522 and BIOS 524; and graduate standing; and consent of the instructor. Corequisites: Requires concurrent registration in BIOS 525.

BIOS 530. Population Ecology. 3 hours.

Life histories, population processes and interactions, and theories of distribution and abundance. Course Information: Prerequisite(s): BIOS 220 and BIOS 221 and BIOS 330 and BIOS 331 and consent of the instructor.

BIOS 531. Introduction to Ecology and Evolution I. 3 hours.

Concepts, techniques, and skills needed for research in ecology and evolution. Course Information: Prerequisite(s): Consent of the instructor.

BIOS 532. Introduction to Ecology and Evolution II. 3 hours.

Evolutionary and physiological research. Course Information: Prerequisite(s): Consent of the instructor.

BIOS 533. Analyzing Ecological Data: Philosophies, Approaches, and Techniques. 4 hours.

Differing philosophies of study design and data analysis in ecological research. Emphasis on the use of the R language for statistical computing to implement a suite of techniques for analyzing univariate and multivariate data. Course Information: Extensive computer use required. Prerequisite(s): BIOS 480; or consent of the instructor.

BIOS 534. Ecology of Biodiversity. 3 hours.

Causes and consequences of different levels of species diversity across a broad spectrum of systems. Core approaches and concepts of community ecology. Application of theory and empirical findings to conserving, managing and restoring biodiversity. Course Information: Prerequisite(s): Consent of the instructor. Recommended Background: Undergraduate courses in basic ecology, calculus, and introductory statistics; at least one graduate course from the following: BIOS 530, BIOS 535, CME 521, UPP 554, or similar courses at UIC or other institutions.

BIOS 539. Seminar in Ecology and Evolution. 0-1 hours.

Graduate student and faculty seminars on selected topics in ecology and evolution. Credit is given only upon completion of a seminar presentation. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated.

BIOS 548. Capstone Project in Landscape, Ecological and Anthropogenic Processes. 4 hours.

Interdisciplinary capstone project course that explores a "real-world" environmental issue selected by the students and approved by the faculty. Students will conduct research and analysis collaboratively and develop solutions and recommendations. Course Information: Same as CME 548 and EAES 548. Prerequisite(s): Grade of B or better in BIOS 540 or Grade of B or better in CME 540 or Grade of B or better in EAES 540 or Grade of B or better in UPP 555; and Grade of B or better in BIOS 546 or Grade of B or better in CME 546 or Grade of B or better in EAES 546 or Grade of B or better in UPP 555; and Grade of B or better in BIOS 547 or Grade of B or better in CME 547 or Grade of B or better in EAES 547 or Grade of B or better in UPP 555. Class Schedule Information: To be properly registered, students must enroll in one Lecture-Discussion and one Studio.

BIOS 552. Chemical Biology. 4 hours.

Major trends and recent developments in research at the interface of chemistry and biology. Course Information: Same as CHEM 552.

BIOS 559. Special Topics in Biochemistry. 3-4 hours.

Selected topics of current interest in biochemistry. Course Information: Same as CHEM 559. May be repeated. Students may register in more than one section per term. Prerequisite(s): CHEM 454 or BIOS 454 or consent of the instructor.

BIOS 560. Topics in Paleontology. 3-4 hours.

In-depth analysis of current problems and issues in paleontology, involving reading primary literature, student presentations, and critical discussions. Course Information: Same as EAES 560. May be repeated if topics vary. Prerequisite(s): Consent of the instructor.

BIOS 565. Modern Methods in Microscopy and Bioimaging. 2 hours.

A graduate-level course surveying modern microscopy and bioimaging methods. Covers both basic principles and practices of major microscopy techniques as well as state-of-the-art literature in the field.

BIOS 582. Methods in Modern Neuroscience. 2 hours.

Underlying principles and applications of techniques used to analyze nervous system organization and function. Behavioral, electrophysiological, anatomical, and biochemical approaches are considered. Course Information: Same as NEUS 582. Animals used in instruction.

BIOS 584. Foundations of Neuroscience I. 3 hours.

Provides a core understanding of modern neuroscience. Focuses on topics in cell and molecular neuroscience. Taught by faculty from multiple units. Course Information: Same as NEUS 501. Recommended background: Credit or concurrent registration in GCLS 503.

BIOS 585. Foundations of Neuroscience II. 3 hours.

A core understanding of modern neuroscience. Focus is on topics in systems, cognitive and behavioral neuroscience. Will be taught by faculty from multiple units. Continuation of NEUS 501. Course Information: Same as NEUS 502. Prerequisite(s): NEUS 501 or BIOS 584. Recommended background: Credit or concurrent registration in NEUS 403.

BIOS 586. Cell and Molecular Neurobiology. 3 hours.

Structure and function of voltage-dependent and neurotransmitter-gated ion channels; the role of these ion channels in synaptic transmission, synaptic modification, and neuromodulation. Course Information: Same as ANAT 586. Prerequisite(s): BIOS 442 or consent of the instructor.

BIOS 592. Research Seminar. 1-2 hours.

Presentation of student research with an emphasis on problem-solving and theoretical implications. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Prerequisite(s): Consent of instructor.

BIOS 593. Introduction to Laboratory Research. 2-6 hours.

A hands-on, in-depth introduction to selected research topics and laboratory techniques designed for the beginning graduate student. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Students may register in more than one section per term. Prerequisite(s): Consent of the instructor.

BIOS 594. Special Topics in Biological Sciences. 1-2 hours.

Selected aspects in biological sciences. Credit varies according to the seminar offered. Course Information: May be repeated. Students may register in more than one section per term.

BIOS 595. Departmental Seminar. 0 hours.

Weekly seminar by staff and invited speakers. Required of graduate students every semester. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated.

BIOS 597. Project Research. 2-8 hours.

Guided research projects on selected topics in specific fields of advanced modern biology. Not to be used for thesis research. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Students may register in more than one section per term. Prerequisite(s): Consent of the instructor.

BIOS 598. Master's Thesis Research. 0-16 hours.

Independent research in specialized projects under the direction of a faculty member with appropriate graduate standing, leading to completion of master's thesis. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Prerequisite(s): Consent of the instructor.

BIOS 599. Doctoral Thesis Research. 0-16 hours.

Independent research on specialized topics under the direction of a faculty member with appropriate graduate standing, leading to completion of Ph.D. thesis. Course Information: Satisfactory/Unsatisfactory grading only. May be repeated. Prerequisite(s): Consent of the instructor.